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SRX7549546: GSM4262903: SacCer.RepTiming.60min.Replicate1a; Saccharomyces cerevisiae; OTHER
1 ILLUMINA (Illumina HiSeq 4000) run: 6.3M spots, 1.9G bases, 718.7Mb downloads

Submitted by: NCBI (GEO)
Study: Isolation and analysis of rereplicated DNA by Rerep-seq
show Abstracthide Abstract
Changes in gene copy number contribute to genomic instability, the onset and progression of cancer, developmental abnormalities, and adaptive potential. The origins of gene amplifications have remained elusive; however, one model suggests DNA rereplication could be a source of gene amplifications. The inability to determine which sequences are rereplicated and under what conditions have made it difficult to determine the validity of these models. Here we present RerepSeq, a technique that selectively fragments and enriches rereplicated DNA in preparation for next generation sequencing. RerepSeq is applicable to any species and can be utilized with low amounts of input DNA with rapid results. We validated RerepSeq by simulating DNA rereplication in yeast and human cells. Using RerepSeq, we demonstrate that rereplication induced in S. Cerevisiae by deregulated origin licensing is non-random and defined by broad domains that span multiple replication origins and topological boundaries. Overall design: Rerep-seq leverages the semiconservative nature of DNA replication to selectively fragment and enrich rereplicated DNA. Cells are labeled with the thymidine analog BrdU for one cell cycle to allow BrdU incorporation into the newly replicated DNA strand, rereplicated regions incorporate BrdU in both strands. Genomic DNA from cells labeled with BrdU is purified and subjected to biochemical processing to induce ssDNA breaks at the sites of BrdU incorporation. This is performed by subjecting DNA to UVA treatment in the presence of Hoechst 33258 to photolyze the bromine from BrdU leaving deoxyuracil. Deoxyuracil is then removed by treatment with UDG (uracil DNA glycosylase) leaving an abasic site. The abasic site is then converted to a single strand DNA break by deoxyribose excision by APE1 (apurinic/apyrimidinic endodeoxyribonuclease). This results in nicked genomic DNA for normally replicated DNA, but staggered nicks producing dsDNA breaks in rereplicated DNA. The fragmented rereplicated DNA can then be isolated by size fractionation and analyzed by quantitative PCR or next-generation sequencing.
Sample: SacCer.RepTiming.60min.Replicate1a
SAMN13838246 • SRS5987066 • All experiments • All runs
Library:
Instrument: Illumina HiSeq 4000
Strategy: OTHER
Source: GENOMIC
Selection: other
Layout: PAIRED
Construction protocol: DNA extraction: As RerepSeq selectively fragments rereplicated DNA and enriches those sequences by size selection, it is imperative that the genomic input DNA is of high molecular weight. For yeast samples: cells were pelleted by centrifugation at 3000 rpm for 3 minutes and resuspended in SCE (1M sorbitol, 100 mM sodium citrate, 10 mM EDTA pH 8.0); fresh 0.125% (v/v) β-mercaptoethanol and 10 U/mL zymolyase was added and incubated for 30-60 minutes at 37ºC to digest cell walls; human cells or yeast spheroplasts were pelleted by centrifugation at 3000 rpm for 3 minutes and resuspended in 500uL RIPA buffer (10 mM Tris-Cl pH 8.0, 1 mM EDTA, 1% Triton X-100, 0.1% sodium deoxycholate, 0.1% SDS, 140 mM NaCl, 1 mM PMSF) with RNase A, 0.2 mg/mL and incubated at 37ºC for 1 hour. 25uL 20% SDS and 10µl of 20 mg/ml proteinase K was then added and allowed in incubate at 55ºC for 2 hours. DNA was extracted twice using Phenol:Chloroform:Isoamyl alcohol (25:24:1, v/v) and precipitated using isopropanol. DNA was resuspended and stored in nuclease free water. RerepSeq Digestion: In an 8 strip 200 µl PCR tube 1 – 5 µg (depending on the experiment) high molecular weight genomic DNA was mixed with 2.5uL 10X Hoechst 33258 (0.1mg/ml) and 2.5uL 10X NEB Buffer 4 (50 mM Potassium Acetate, 20 mM Tris-acetate , 10 mM Magnesium Acetate, 1 mM DTT pH 7.9@25°C) to a final volume of 24uL; open tubes placed upright in PCR tube rack, covered with glass plate (3" x 3" glass plate from VWR Vertical Gel Electrophoresis Systems), exposed to 7.5 minutes of glass plate filtered (UVA only) from a Stratalinker. Following UVA treatment, samples were digested with 0.5uL UDG (5 units of Uracil-DNA Glycosylase), 0.5uL APE1 (10 units of human apurinic/apyrimidinic endonuclease 1) for 2 hours at 37C. Digested DNA was repaired with NEB's FFPE DNA Repair Mix for 30 minutes then separated on 0.8% agarose gel for 15 minutes. Fragmented DNA ranging from 0.1Kb to 3Kb was gel extracted with Wizard® SV Gel and PCR Clean-Up System (Promega) and the purified DNA was resuspended in 50uL of water for subsequent qPCR and sequencing analysis. Illumina Nextera DNA Flex Library Prep Kit
Experiment attributes:
GEO Accession: GSM4262903
Links:
Runs: 1 run, 6.3M spots, 1.9G bases, 718.7Mb
Run# of Spots# of BasesSizePublished
SRR108804326,336,9511.9G718.7Mb2020-01-23

ID:
9863619

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